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The air explodes with the sound of high-powered rifles and the startled infant watches his family fall to the ground, the image seared into his memory. He and other orphans are then transported to distant locales to start new lives. Ten years later, the teenaged orphans begin a killing rampage, leaving more than a hundred victims.

A scene describing post-traumatic stress disorder (PTSD) in Kosovo or Rwanda? The similarities are striking — but here, the teenagers are young elephants and the victims, rhinoceroses. In the past, animal studies have been used to make inferences about human behavior. Now studies of human PTSD can be instructive in understanding how violence also affects elephant culture.

Psychobiological trauma in humans is increasingly encountered as a legacy of war and socio- ecological disruptions. Trauma affects society directly through an individual’s experience, and indirectly through social transmission and the collapse of traditional social structures. Long-term studies show that although many individuals survive, they may face a lifelong struggle with depression, suicide or behavioural dysfunctions. In addition, their children and families can exhibit similar symptoms, including domestic violence. Trauma can define a culture.

How PTSD manifests has long been a puzzle, but researchers today have a better idea as to why the effects of violence persist so long after the event. Studies on animals and human genocide survivors indicate that trauma early in life has lasting psychophysiological effects on brain and behavior.

Under normal conditions, early mother-infant interactions facilitate the development of self-regulatory structures located in the corticolimbic region of the brain’s right hemisphere. But with trauma, an enduring right-brain dysfunction can develop, creating a vulnerability to PTSD and a predisposition to violence in adulthood. Profound disruptions to the attachment bonding process, such as maternal separation, deprivation or trauma, can upset psychobiological and neurochemical regulation in the developing brain, leading to abnormal neurogenesis, synaptogenesis and neurochemical differentiation. The absence of compensatory social structures, such as older generations, can also impede recovery.

Elephant society in Africa has been decimated by mass deaths and social breakdown from poaching, culls and habitat loss. From an estimated ten million elephants in the early 1900s, there are only half a million left today. Wild elephants are displaying symptoms associated with human PTSD: abnormal startle response, depression, unpredictable asocial behavior and hyperaggression.

Elephants are renowned for their close relationships. Young elephants are reared in a matriarchal society, embedded in complex layers of extended family. Culls and illegal poaching have fragmented these patterns of social attachment by eliminating the supportive stratum of the matriarch and older female caretakers.

Calves witnessing culls and those raised by young, inexperienced mothers are high-risk candidates for later disorders, including an inability to regulate stress-reactive aggressive states. Even the fetuses of young pregnant females can be affected by pre-natal stress during culls. The rhinoceros- killing males may have been particularly vulnerable to the effects of pre-and postnatal stress for two reasons. Studies on a variety of species indicated that male mammalian brains develop at a slower rate relative to females, but also that elephant males require a second distinct phase of socialization.

Elephant hyperaggression is not an isolated event. At another heavily affected African park, intraspecific mortality among male elephants accounts for nearly 90% of all male deaths, compared with 6% in relatively unstressed communities. Elsewhere, including Asia, there are reports of poor mothering skills, infant rejection, increased problem animals and elevated stress-hormone levels.

Neuroscience has demonstrated that all mammals share a ubiquitous developmental attachment mechanism and a common stress-regulating neurophysiology. Now, a wealth of human-animal studies and the experience of human victims of violence are available to help elephants and other species survive.

单选题 According to the essay, elephant hyperaggression _____.
【正确答案】 C
【答案解析】根据文章第二段可知“Now studies of human PTSD can be instructive in understanding how violence also affects elephant culture”, 现在对人类创伤后应激性障碍( PTSD) 的研究可以对理解暴力如何影响大象文化起指导作用。 后文都是在这个基础上展开论述的, 因此, 根据文章大象的高度侵略性跟其经受过的心理创伤有关, 故选C。
单选题 Which of the following statements about post-traumatic stress disorder that elephants suffer from is NOT true?
【正确答案】 D
【答案解析】根据文章第八段可知“Calves witnessing culls and those raised by young, inexperienced mothers are high-risk candidates for later disorders, including an inability to regulate stress-reactive aggressive states. Even the fetuses of young pregnant females can be affected by pre-natal stress during culls”, 即: 目睹过宰杀过程的小象以及由年轻的、 没有经验的雌象哺育的小象是晚期疾病的高发群体, 包括无法调节应激下的侵略性状态。 宰杀期间年轻雌象肚里的胎儿甚至也会受到产前压力的影响。 因此, 造成大象创伤后应激性障碍的原因是曾经目睹过同类被宰杀或者是雌象产前面临巨大压力。 故D选项表述错误。 根据上一题可知选项A、 B、 C的表述均符合原文, 故选D。
单选题 Which one of the following statements about elephants is NOT correct?
【正确答案】 D
【答案解析】根据文章第七段可知“Elephants are renowned for their close relationships. Young elephants are reared in a matriarchal society, embedded in complex layers of extended family. Culls and illegal poaching have fragmented these patterns of social attachment by eliminating the supportive stratum of the matriarch and older female caretakers”, 大象以其同类间的亲密关系而著称, 年轻的大象是在母系社会中长大的, 是复杂大家庭中的一员。 宰杀和非法偷猎雌象和年老的负责照顾后代的雌象使这种社会的依附模式支离破碎。 因此选项A、 B、 C均表述正确, D选项文中未提及, 故选D。
单选题 Which one is NOT the symptom of post-traumatic stress disorder of elephants?
【正确答案】 B
【答案解析】根据文章第六段可知“Wild elephants are displaying symptoms associated with human PTSD: abnormal startle response, depression, unpredictable asocial behavior and hyperaggression”, 野生大象在受过创伤后会呈现出和人类创伤后应激性障碍有关的症状: 异常惊吓反应、 抑郁、 不可预测的孤僻行为和高度侵略性。 因此A、 C、 D均符合文意。 B( 反复无常的行为) 文中未提及, 故选B。
单选题 According to the essay, why do the effects of violence that might cause the breaking bond between the mother and the infant persist so long after the event?
【正确答案】 B
【答案解析】根据文章第五段可知“Under normal conditions, early mother-infant interactions facilitate the development of self-regulatory structures located in the corticolimbic region of the brain’s right hemisphere. But with trauma, an enduring right-brain dysfunction can develop, creating a vulnerability to PTSD and a predisposition to violence in adulthood. Profound disruptions to the attachment bonding process, such as maternal separation, deprivation or trauma, can upset psychobiological and neurochemical regulation in the developing brain, leading to abnormal neurogenesis, synaptogenesis and neurochemical differentiation. The absence of compensatory social structures, such as older generations, can also impede recovery”, 在正常情况下, 雌象与小象之 间早期的互动促进了位于大脑右半球皮质边缘地区的自我调节构造的发展。 但由于受到创伤, 右脑功能障碍持续出现, 造成成年大象创伤后精神障碍症和易患性暴力倾向。 对依附性 亲密过程的严重破坏, 如雌象被迫分离、 掠夺或创伤, 可以扰乱大脑发育中的精神生物学和 神经化学调节, 导致神经、 突触形成和神经化学的分化。 社会结构补偿的缺失, 如老一代的 大象, 也可能阻碍创伤的恢复。 只有B选项符合文意, 故选B。 
单选题 What does the underlined word “ubiquitous” in the last paragraph possibly mean?
【正确答案】 B
【答案解析】分析文章最后一段可知: 神经科学已经证明, 所有哺乳动物都有一种…的发育依恋机制和一种共同的应激调节的神经生理学。 现在, 大量的人类动物研究和暴力受害者的经验可以帮助大象和其他物种在创伤中生存下来。 “ubiquitous”和后文的common相呼应, 都表明哺乳动物之间的共同点, 只有universal( 通用的, 普遍的) 最符合题意, 故选B。