单选题. ①Noting that bison herds appear to graze selectively in areas occupied by prairie dog colonies, Coppock hypothesized that the attraction is due to improved forage quality: prairie dogs' own grazing activities alter herbage dynamics, plant species composition, and nutrient cycling. ②The colonies' territory contains relatively little low-quality mature standing herbage and is characterized by communities of plants that are high in crude protein and highly digestible. ③However, such effects are not produced only by prairie dogs. ④Willms showed that sites selectively grazed by cattle had reduced standing dead plant material, altered species composition, and increased soil nitrates, ammonium, and available phosphorus. ⑤In fact, long-term, intensive use by any grazer will cause comparable changes in plant communities.29. The highlighted sentence serves primarily to ______