摘要
大黄鱼的精子由头部和尾部（鞭毛）两部分组成。头部结构较为独特。其腹侧有一较大的细胞核，背部有中心粒复合体。头部的后端是袖套。细胞核的腹面稍向外突出，背面则稍向内凹。细胞核中的染色质浓缩成致密的团块状。团块状的染色质之间分布着松散的纤维状染色质。植入窝位于细胞核的背部表面，由细胞核背面向内凹陷而成，呈一沟状，其走向与精子的长轴平行。中心粒复合体位于植入窝内。近端中心粒的长轴与精子的长轴垂直，基体的长轴与精子的长轴平行。基体的头端呈筒状，由电子致密物质构成，其间分布着少量的微管。基体的尾端，筒状结构分裂成九束。袖套中分布着线粒体和一些囊泡。近袖套内膜的细胞质中存在着一层与袖套内膜平行并且比它薄的膜。尾部从袖套腔中伸出。尾部细长，分为主段和末段。主段的中心结构是轴丝。轴丝外仅有少许细胞质。轴丝与基体的尾端相连。轴丝起始端的结构较为特殊。该处无中央微管对，九组外周二联微管仅具雏型，Ａ小管和Ｂ小管的管腔呈电子致密状，也无动力蛋白臂附于二联微管之上。在主段的末端，构成轴丝的微管螺旋成篓状结构。一小部分微管从篓状结构延伸进入末段。
The spermatozoon of the teleost, Pseudosciaena crocea (Richardson) consists of two parts: the head and the tail (flagellum). The structure of the head is quite special. A large nucleus is located at the ventral part of the head. The dorsal part of the head is occupied by the centriolar complex. The posterior end of the head is the sleeve. The ventral surface of the nucleus is convex,whereas the dorsal surface of the nucleus is concave.Within the nucleus the chromatin condenses into wads,among which are loose chromatin fibers. The implantation fossa is on the dorsal surface of the nucleus. It is groove like running from the anterior end to the posterior end of the nucleus. The centriolar complex is in the implantation fossa. The proximal centriole is perpendicular to the major axis of the spermatozoon,whereas the basal body is parallel to the major axis of the spermatozoon. The anterior end of the basal body is roll like. It is electron dense, with some microtubules in it. The posterior end of the basal body splits into 9 strips. The sleeve contains several mitochondria and vesicles. Within the cytoplasm of the sleeve, there is a membrane which is near and parallel to, as well as thinner than the inner membrane of the sleeve. The tail, which is thin and long, stretches out of the central space of the sleeve. It consists of a principal piece and an end piece. The central structure of the principal piece is axoneme. There is only a little cytoplasm outside the axoneme. The 9 doublets of the axoneme is connected to the 9 strips of the posterior end of the basal body. The initial part of the axoneme is special in structure: there is no central pair of the single microtubule, the doublets of it are electron dense, and there are no dynein arms on the doublets. At the end of the principal piece, the microtubules, which constitute the axoneme, spiral to become a basket like structure.Several microtubules extend into the end piece of the tail.
出处
《动物学报》
SCIE
CAS
CSCD
1997年第2期119-126,共8页
ACTA ZOOLOGICA SINICA
关键词
鱼纲
大黄鱼
精子
超微结构
Pseudosciaena crocea(Richardson),Spermatozoon,Ultrastructure, Implantation fossa, Centriolar complex,Basket like structure
