摘要
太白红杉 (LarixchinensisBeissn)的雄球花 7月初开始分化。小孢子囊壁一般包括 5～ 6层细胞 :表皮、药室内壁、2～ 3层中层和绒毡层。绒毡层属于周原质团型。造孢细胞在 7月下旬形成 ,8月上旬形成小孢子母细胞 ,8月下旬开始减数分裂 ,于 1 0月上旬进入双线期 ,并以双线期渡过休眠。翌年 3月下旬解除休眠继续进行减数分裂 ,4月中旬形成四分体 ,4月下旬到 5月初小孢子从四分体内释放出来 ,小孢子经过连续 4次有丝分裂后 ,于 5月中旬形成 5_细胞型的成熟花粉粒 (雄配子体 )并开始散粉。小孢子母细胞发育表现出不同步现象 ,部分小孢子母细胞在发育过程中出现退化 。
The male cone of Larix chinensis Beissn differentiates at the beginning of July. The microsporangium wall of L. chinensis comprises one epidermis, one endothecium, 2～3 middle layers, and a tapetum. The tapetum conforms to periplasmodial type. The sporogenous cell occurs in late July. The microspore mother cell is formed in early August, and its meiosis begins to occur in late August. The microspore mother cell enters into diplotene in early October, and goes through dormancy at this phase. The microspore mother cell ends dormancy in late March next year, and continues its meiosis. In the middle of April tetrad forms and microspore is released from late April to early May. After microspore’s four times continuous mitosis, the mature pollen (male gametophyte) which comprising five cells forms and begins shedding in middle May. The development of the microspore mother cell is not synchronous, and some microsporocytes begin to degenerate, resulting in the formation of a large cavity in the microsporangium.
出处
《植物学通报》
CSCD
北大核心
2003年第5期576-584,共9页
Chinese Bulletin of Botany
基金
陕西省教育厅重点科研计划资助项目 (0 0JK0 0 7)
关键词
太白红杉
小孢子母细胞
雄配子体
发育
松科
Larix chinensis Beissn, Microspore mother cell, Male gametophyte, Development
