摘要
本文对春兰(Cymabidium goeringii(Rchb.f.)Rchb.f.)花药壁发育的两个时期进行了光镜和电镜的观察。阐明了花药壁的解剖学及其各种组织的细胞学特点。绒毡层是独特的,有如下特点:(1)具内绒毡层,它与外绒毡层成为一个连续组织包围着花粉块。内绒毡层是由连结组织衍生。(2)内外绒毡层都是由2—3层的细胞组成,具一致的超微结构特点和表现相同的功能。(3)在较早的发育时期,绒毡层细胞中存在许多的含脂质体,这是后期绒毡层细胞贮存大量脂体的主要来源。这种含脂质体是由核被膜的扩展产生。(4)在四合花粉形成后,绒毡层细胞的分泌物及细胞自溶后的产物,不仅沉积在花粉块的表面形成覆盖层,而且流入花粉块的内部四合花粉之间,表现为加强四合花粉之间的粘合的作用。本研究结果对被子植物绒毡层的结构和功能提供了一些新的资料。
The structure of the anther wall in Cymbidum goeringii (Rchb. f.)Rchb.f, was followed through two developmental stages using light and electron microscope. The peculiar anatomicaI pattern of the anther wall and the cytological characteristics of its tissues are described. Tapetam is the most notable feature in the anther wall. The specialized characteristics of the tapetal cells may be summerized as followings: (1) The tapetum may be distinguished into outer and inner rapetum, but they constitute the continuous layers surrounding the pollinium. The inner tapetum is derived from the connective tissue. (2) The outer and inner tapetum are composed of 2—3 celt layers. They have the same ultrastructural feauares and functions. (3) There are many lipid-con- taing plastids in the tapetal cells during theii developmental stage. This is the main source of lipid bodies in the aapetal cells during their fina|developmental stage. The lipid-containing pla- slids are derived from the extension of the nuclear membrane. (4) After the formation of te- trads, the secretion and the autolysates from of the tapetal cells not only deposition the surface of the pollinium to form the film, but also flow intn the inner region of the polinia between the te- lads contributing to their adhesion. The resul~,s of this study provide some new data of the ia- petal structure and function in ^+he angiosperms.
关键词
春兰
药室内壁
绒毡层
脂类合成
Cymbidium goeringii
Andothecium
Tapetum
Lipid synthesis
