基金The project was supported by the National Natural Science Foundation of China(Grant Nos.1157119111431006+3 种基金11371202)the Scientific Research Fund of Liaoning Provincial Education Department of China(Grant No.LSNQN201912)the Science and Technology Project of Hebei Science and Technology Department(Grant No.162176489)the Education and Teaching Research Project of Shenyang Agricultural University(Grant No.2018-84)