基金Supported by the National Natural Science Foundation of China(30660120)the Program of Science and Technology Department of Qinghai Province(2010-H-809)+2 种基金the Science and Technology Key Project of the Ministry of Education(209133)the National Basic Research Program of China(973 Program)(2009CB421102)the National Major Special Project on New Varieties Cultivation for Transgenic Organisms(2009ZX08009-130B to J.Xu)