基金Supported by National Key Technology Research and Development Program(2012BAD04B01-02)Science Foundation of Heilongjiang Province for Outstanding Young Scientists(JG05-22)+2 种基金Key Science and Technology Program of Heilongjiang Province(GA09B102-3)Youth Foundation for Agricultural Science and Technology Innovation in Heilongjiang Province in 2012Heilongjiang Postdoctoral Sustentation Fund(LBH-Z10038)~~